Pests
Pests have specialized in living in storage rooms, where the abundance of heat, moisture and food produces their mass multiplication. Some damage the field, and after harvesting they increase their population density in the storage rooms (rice weevil, grain moth). There are insects that have adapted to such an extent that their life outside the storage rooms is impossible (the grain weevil has degenerated wings).
Grain moth, dried fruit moth, gray flour moth are the most characteristic storage representatives of the order Lepidoptera (butterflies), damaging in mills, silos, homes. Storage moths fly around and infect only the surface of the stored material, after which they can migrate to other areas.
Nocturnal insects that rarely fly during the day and are often found perched on walls. Adult moths do not feed, but break down the nutrients in their bodies accumulated during the caterpillar stage.
The gray flour moth also forms cobwebs that clog elevators and augers in mills, and its larvae gnaw through sieves. Warehouse enemies.
Wheat and rice weevils, as well as the Surinamese mealworm and the Moorish beetle are representatives of the order Coleoptera (beetles), damaging granaries, bakeries, mills, and candy and pasta warehouses. Warehouse beetles enter and infect the interior of the stored grain layer, and when the grain warms up, they migrate in the direction of the higher temperature. The larvae gnaw through the interior of the grain. Weevils attack only whole grains and to prevent their activity it is necessary for the grain to be ground (feed). They transmit Escherichia coli, as well as aflatoxin through the mold Aspergillus flavus.
The wheat and rice weevils occur together and contaminate grain feeds, making them unsuitable and dangerous for animal food. The Surinamese mealworm multiplies in broken grains and grains already damaged by proboscis and moth larvae. The adult Moorish beetle feeds on grain, flour, pasta, gnaws at mill screens and cardboard packaging, and its larva gnaws at the inside of the husked rice.
Harm
Storage pests cause enormous losses to the country’s economy, due to their high fertility, rapid reproduction and the lack of any compensatory mechanisms for damaged food and materials. Therefore, there is no Threshold of Economic Harm for them.
The damage caused by pests to stored grain is no less than that caused by pests in the field, but in stored products they most often remain hidden, at least at the beginning of their harmful activity, while the density is relatively low.
The appetite of some storage pests is impressive:
– an adult grain weevil eats about 1 g of grain during its life (the beetle weighs less than 1 milligram);
– a rat eats 22 kg of grain every year;
– a house sparrow pecks 12 – 13 g of grain per day.
Storage pests contaminate products and materials with::
– eggs (chorions);
– larvae, pupae and their body shells;
– excrement and corpses of dead adults.
Storage pests also cause indirect damage. They cause self-heating of the bulk grain during their active activity. In the foci of self-heating, the temperature rises to 42°C, with a grain humidity of 15%. During self-heating, the surface of the grain layer becomes moist and a crust forms. This phenomenon is best observed in autumn, when the surface of the bulk grain cools faster. Water vapor moves from the place of self-heating to the cool surface layer, where it causes mold and seed germination to a depth of 15-20 cm. Sometimes self-heating is of even higher value and can lead to self-ignition.








The flat grain beetle (Cryptolestes testaceus) reaches sizes up to 2.5 mm. The body is rusty in color, densely covered with hairs. The beetles have wings and fly well. The beetles live up to six months, they can survive without food for 2.5 months. The female lays several dozen eggs. After a few days, larvae hatch from the eggs. The development of one generation lasts from 70 to 100 days. During the year, 3 to 5 generations can develop.
They do not tolerate low humidity. They develop best at 20-23 °C and in an environment with high air humidity. Their development stops at a temperature of 12-14 °C.



The gray flour moth (Ephestia kuehniella) develops 3-4 generations per year in unheated warehouses, and in heated premises it can develop year-round and have up to 8-10 generations. It overwinters as a caterpillar, in a silk cocoon and limitedly as a pupa (in unheated warehouses). Usually in the second half of April – the first half of May, the caterpillars pupate. The pupal stage lasts 5-7 days.
The imagined butterflies are sexually mature and shortly after imagining they copulate. They are active at night, and during the day they hide in dark places. One female lays 200-300 eggs, singly or in small clusters. The eggs are laid in cracks in the walls, on the floor, sacks, food and other places. At a temperature of 13-150C, the egg stage lasts 12-21 days, and at 22-240C – 6-10 days.The hatched caterpillars crawl to the stored products and begin to feed by gnawing them. They secrete abundant silk threads, with which they bind the food products in the form of large balls (globes), reaching from half to several kilograms. With these balls they can clog the sieves and elevators in the mills and large warehouses. The caterpillars have extremely strong jaws and are able to gnaw through the thin sieves in the mills, which may require their replacement and stopping the work process. The duration of their development depends mainly on the temperature. When it is favorable, it ends in 17-20 days. The food host is also important.
The fed caterpillars leave the feeding areas and go into cracks in the floor and walls, in the attics, or to other hiding places, where they spin a silk cocoon and pupate in it.

The tobacco beetle (Lasioderma serricorne) develops 3 generations per year and overwinters as larvae of different ages in the damaged areas. Younger larvae usually die in winter or early spring, as they are sensitive to low temperatures. In heated warehouses, the tobacco beetle develops year-round. When the temperature rises above 10-120C, the immature larvae fully develop and pupate in a special chamber, in the folded parts of cigarette and cigar boxes, inside the damaged tobacco, along the main vein of raw tobacco leaves and in other protected places..

The Surinamese flour beetle (Oryzaephilus surinamensis) is a small beetle with a body length of about 2–2.5 mm. The female lays up to 400 eggs on cereals and grains, flour (hence its name), dried fruits and spices. The larvae develop rapidly, especially in damp places (with a relative humidity of more than 14%), feeding on stored products. They pupate in cracks and crevices. The larvae complete their development in 4–6 weeks, depending on the air temperature. Adults sometimes live for a long time, up to 3 years.

American cockroaches, (Periplaneta americana) reach a length of 34-53 mm. They are reddish brown in color, with a pale brown to yellowish band around the edge of their pronotal shield. Both male and female American cockroaches have wings, but their flight is very poorly developed. They inhabit a wide variety of environments, and can live in very dry climates, as long as they have access to water. They prefer temperatures of 29 degrees and cannot tolerate cold temperatures.

The gray rat (Rattus norvegicus), also known as the barn rat, has a relatively long body up to 25 cm. The tail is slightly shorter or equal to the body length of 22–25 cm. They give birth 2-4 times a year, 6-10 to 17 young.
Newborns are blind and see after two weeks. The mother nurses for one month. After 3–4 months, they reach sexual maturity. They live up to 3 years. They feed on both plant and animal food.

The grain beetle (Rhyzopertha dominica) is a primary enemy of cereals and seeds. It destroys almost completely the grains, leaving only part of the husks. The larvae destroy about 30% of the endosperm of the grain. In one day, the beetles destroy an amount of grain that is approximately equal to their mass. With a large population of grain beetles, the grain acquires a characteristic musty honey smell.

Ticks (Ixodoidea) are a superfamily of small arachnids of the subclass Acari. They are ectadartisiophorasites that feed by sucking blood from mammals, birds, and in some cases reptiles and amphibians. They are an important vector for the spread of many infectious diseases. The true parasite is the female tick. After fertilization, it digs small tunnels /galleries/ in the uppermost layer of the skin, in which it lays 24 eggs measuring 0.10-0.16 mm. By the third to seventh day, they turn into larvae, and by the 17th day – into nymphs, which are already capable of transmitting the infection. Blood-sucking ticks inhabit grassy-shrubby and forest areas and prefer high humidity.

The rusty flour beetle (Tribolium confusum) has an elongated body up to 3.5 mm long, with a shiny rusty-red color. It has well-developed wings and flies well, especially at night.
The optimal temperature for the development of the rusty-red flour beetle is about 30°C. At this temperature, the female can lay an average of 300-400 eggs and a maximum of 1000. The species can also develop at a grain humidity of 11-12%. Under optimal conditions, the life cycle from egg to adult takes about 32-35 days and can develop from 6 to 8 generations per year. Adults live up to two years.

Trogoderma (Trogoderma granarium) is one of the most dangerous species attacking grain products. It damages the seeds of wheat, barley, corn, rice, peanuts, cotton, legumes and others. Among the damaged products there is a large amount of larval cuticles. The fastest development occurs at 30 – 35 C, and the lower threshold is 8 C. At low temperature and low humidity, the larvae hide in cracks and fall into diapause and withstand hunger for up to 4 years.